FERN PLANTS GALLERY PAGES Fern Culture with British Ferns and their Allies comprising the Ferns, Club-mosses, Pepperworts and Horsetails by Thomas Moore, F.L.S, F.H.S., Etc. London George Routledge and Sons, Broadway, Ludgate Hill. Hardcover published in 1861 provides details on British Ferns |
TYPE OF FERN TO GROW
Where to see UNITED STATES WALES |
USE OF FERN
Where to see AUSTRALIA CANADA ENGLAND FRANCE GERMANY IRELAND NETHERLANDS
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SPORE COLOUR BED PICTURES Where to see NEW ZEALAND SCOTLAND UNITED STATES |
A Natural History of Britain's Ferns by Christopher N. Page. Published by William Collins Sons & Co. Ltd in 1988. ISBN 0 00 219382 5 (limpback edition) provides details of Coastal, Man-made Landscapes, Woodland, Wetland, Grassland and Rock Outcrops, Heath and Moorland, Lower Mountain Habitats, Upper Mountain Habitats and Atlantic Fringe Ferns. Tree Ferns by Mark F. Large & John E. Braggins. Published by Timber Press in 2004. ISBN 978-1-60469-176-4 is a scientifically accurate book dealing with Tree Fern species cultivated in the United States and the Pacific, but little known and rare tree ferns are also included. The Observer's Book of Ferns, revised by Francis Rose, previous editions compiled by W.J.Stokoe. Published by Frederick Warne & Co. Ltd in 1965 provides a comprehensive guide to 45 British species of Ferns. It provides details of habitat and how to use those ferns. The Plant Lover's Guide to Ferns by Richard Steffen & Sue Olsen. Published in 2015 by Timber Press, Inc. ISBN 978-1-60469- Success with Indoor Ferns, edited by Lesley Young. Reprinted 1998. ISBN 1 85391 554 8. It details the care of indoor ferns with their position, choice and fern care. |
Where to see UNITED STATES |
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Fern |
Foliage Colour and |
Height x Spread in inches (cms) |
Type of Fern to Grow |
Use of Fern |
Comments |
Frond |
Credit |
Form |
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Blechnum penna-marina |
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Blechnum spicant |
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Dennstaedtia punctilobula |
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Dryopteris clintoniana |
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Dryopteris expansa |
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Dryopteris tokyoensis |
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Osmunda claytoniana Interrupted Fern, Flowering Fern Very hardy, Native to northeastern North America, India and Asia. The unusual common name for this fern arises because on the fertile fronds the fertile segments are carried in between sets of normal barren segments, giving the appearance of a gap in the frond. Young fronds are covered with wooly, pinkish hairs. |
The leaves grow from a rhizome growing at or below the ground. Forming a lovely spreading vase habit, this low-maintenance native fern makes a distinctive addition to the shade border or woodland garden. |
24-36 x 24-36 |
Primitive Ferns and Fern Oddities Hardy deciduous fern. |
Suitable for Accent Fern. Ferns for Acid Soils. Evergreen and Deciduous Ferns. Ferns for Wet Soils. Cold-hardy Ferns. Rock Garden and Wall Ferns. Shade-Tolerant Fern.
Grows well with hostas in shaded woodland or wild gardens. Also effective along ponds or streams. Interesting accent for the shaded border. |
This clump-forming fern has erect rhizomes that form occasional offshoots and grows in moist-wet to wet, acidic garden soil. The plants have deciduous fronds and do poorly in the Gulf States and subtropical climates. Habitat in forests, shores of rivers or lakes, swamps, wetland margins (edges of wetlands). Easily grown in medium to wet soils in part shade to full shade. Prefers moist, rich, humusy, acidic soils, but adapts to lesser conditions. Deer resistant. |
Osmunda claytoniana.
Interrupted fern, Osmunda claytoniana, in Lac-Mégantic, Quebec. |
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Phegopteris connectilis |
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Polystichum, various species see below |
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Matteuccia struthiopteris Ostrich Fern, Shuttlecock Fern Very hardy. |
Mid Green pinnate fronds taper at both ends and grow in a vase-like cluster around the robust rootstock. A vigorous fern which spreads by underground rhizomes, producng clusters of new fronds at intervals Flushes of new fronds are most decorative. Requires acid, loamy soil. The sterile leaves are deciduous, but the fertile ones persist throughout the winter and shed their spores in early spring, sometimes over snow. Usually 1 or 2 new plants are produced each year from the stolons. |
66 x 36 Spacing 24-30 (60-75) |
Shielder Ferns, Buckler Ferns, Holly Ferns and their Relatives Large hardy deciduous fern. Outdoor Culture: Soil, 2 parts good loam, 1 part leaf-mould. Position, semi-shaded, cool, moist border or margin of pond. Plant, April. |
Suitable for Fern for Acid Soils Cold-hardy Ferns. Border and Foundation Ferns. Ground Cover Ferns. Outdoor Containers. Ferns for Wet Soils |
Grow in moist shade in a woodland garden, a damp border or at the edge of a pond. Mass in moist, shady woodland areas, wild gardens or wet areas near streams or ponds. Combines well with astilbes or hostas. Plant in conjunction with early spring wildflowers (e.g., trilliums, bloodroot, trout lilies or Dutchman's breeches) which will be well on the way toward dormancy by the time this fern reaches full size. It will grow in full sun with constant moisture. Clump-forming fern to grow in Part Shade and Full Shade in Medium to Wet soil. |
Matteuccia struthiopteris, young plant, Hurum, Buskerud (Norway). By Bjoertvedt via Wikimedia Commons.
English: Matteuccia Struthiopteris in Ypäjä, Tavastia Proper, Finland
Русский: Страусник обыкновенный. Спорофиллы и вайи. Россия, Савинский район Ивановской области.
Matteuccia struthiopteris |
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Onoclea Sensitive Fern, Bead Fern, Sympathy Fern Native to North America, Canada and North Asia. Onoclea comes from the Greek onos, vessel, and klein, to close, referring to the pinnules of the fertile leaf, which roll up into bead-like segments to enclose the sori. A member of the Woodsiaceae Cliff Fern Family. Zone 4
Available in USA from ArcheWild Native Nurseries - Quakertown, PA Toadshade Wildflower Farm - Frenchtown, NJ Prairie Nursery - Westfield, WI Yellow Springs Farm Native Plant Nursery - Chester Springs , PA |
Upright, then arching, lance-shaped or triangular, pinnate, pale green sterile fronds. The bead-like appearance of the fertile fronds accounts for this genus's common name of bead fern. Some say that the name sensitive fern originates from the frond's sensitivity to frost (they wither after the first subfreezing temperatures). Winter survival will be enhanced if the dried fronds are left on the plant through the winter. |
36-48 x 36-48 |
Miscellaneous Ferns Hardy deciduous ferns. Fronds, barren ones, broad, once-divided, green; fertile ones, narrow, contracted, once-divided, brown. |
Suitable for Ferns for Acid Soils. Ferns suitable for Outdoor Containers Ferns for Wet Soils Cold-Hardy Ferns The fertile fronds are often used in dried flower arrangements. Best in wet woodland gardens and moist locations alongside streams and ponds. Can grow in very wet soils as long as there is adequate oxygen. It cannot tolerate sour clay or stagnant water. Also, does not tolerate freezing well, turns black even in light frost. Shelters salamanders and frogs |
Open swamps, thickets, marshes, or low woods, in sunny or shaded locations, often forming thick stands from sea level to elevations of 1500 metres. It grows best in a shaded or partially shaded area in a moist soil. The plant can tolerate dryer conditions in shade, and will tolerate wet soils and so occurs in soggy ground or at the very edge of water in shade or sun. Sensitive ferns spread to form colonies and are often the first species to inhabit disturbed areas. They can become weedy if not sited properly. |
Rodgersia and Onoclea. 25 April 2014, 16:50. By peganum from Henfield, England via Wikimedia Commons
日本語: Onoclea sensibilis:コウヤワラビ
Juvenile Onoclea sensibilis sterile fronds in pots. By Coblands. |
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Royal Fern, Flowering Fern Very hardy, |
A fibrous rootstock bears dense clumps of triangular-ovate-pinnate, bright green sterile fronds. In summer, partially fertile fronds, to 6 feet long, have tassel-like tips, with brown or rust-coloured sporangia covering the much smaller pinnae. In autumn, they turn bronze before dying back. This deciduous fern forms a natural, rounded shape and looks fantastic planted near a pond or stream, where its feathery fronds will be reflected in the water. It likes damp, preferably acid soil, and looks breathtaking with other moisture-loving, large foliage plants such as rodgersia and gunnera. |
72 x 144 |
Primitive Ferns and Fern Oddities Hardy deciduous fern. |
Ferns suitable for Outdoor Containers. Ferns for Wet Soils. Bog or Wet-Soil Fern. Evergreen and Deciduous Ferns. Cold-Hardy Ferns. Ferns for Acid Soils. Shade-Tolerant Fern.
Excellent selection for wet areas along ponds, streams, water gardens or in bogs. Also grows well in shaded borders, woodland gardens, wild gardens or native plant gardens. |
Grow in a damp border, or at the margins of a pond or stream. It prefers cool summer climates where it tolerates close to full sun as long as given consistent moisture. Full sun exposure is not recommended for the hot St. Louis summers. |
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Osmunda regalis Image 1 on left from Denver Botanic Gardens
Osmunda regalis Image 2 from Denver Botanic Gardens
Osmunda regalis on right. By Ghislain118 http://www.fleurs-des-montagnes.net via Wikimedia Commons
Nederlands: Plant - Koningsvaren - Osmunda regalis |
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Aglaomorpha coronans Crowning Bear's Claw Fern, Santa Rosa Fern, Basket Fern Semi-tender or hardier Native to Bangladesh, India, China, Taiwan, southeastern Asia, Malaysia, and the Ryuku Islands of Japan. Aglaomorpha is from the Greek aglaos (bright, pleasing) and morphe (shape) |
Deep green ornately lobed outarching "fronds" with sing like basal oak leaf pattern anchored on large sienna pelted showy rhizomes. Very sturdy and reliable.
This plant has been said to grow in the following regions: Clayton, California Hayward, California Long Beach, California Stockton, California Brandon, Florida |
36-48 x Spacing 72-96 (180-240) |
Staghorn Ferns Requires consistently moist soil; do not let dry out between waterings. Always makes an impressive basket fern when well grown. Does best in very couarse, porous epiphyt potting mix and allow it getting fairly dry between watering. "On rather dry or mossy rocks or on tree trunks in open places or in dense forests, not so rare at medium altitudes throughout Thailand." from Ferns of Thailand, Laos and Cambodia |
Suitable for Tropical - in Denver Botanic Garden - Conservatory or Heated Greenhouse Fern for Acid Soil: 5.6-6.0 pH Basket Ferns |
Best grown in well-drained, moist-dry potting mix or uncut moss. The species is easy to grow and thrives in both humid and drier environments. It can be cultivated in the ground - if given sufficient drainage by using a coarse medium - and in hanging baskets. They collect falling leaves and other organic debris that eventually decay to provide a soil for the roots. The dry, papery texture of this humus-collecting leaf-base resists decay. Though large, they are suitable as indoor plants because they tolerate low humidity and irregular watering. |
Glasshouse Works has photos of this fern.
The perennials prefer a shady situation on moist soil. They tolerate temperatures only above at least 1°C (USDA zone 10). Germination Instructions from Exotic Plants. |
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Polystichum andersonii Hardy Species in Zone 5(6) Native to North America, Pacific Northwest. Found growing in the mountains from Alaska south to Oregon and east to Montana. In Zones 6-9. |
Ascending to erect rhizomes and evergreen, bud-bearing leaves - see Section 9 - Propagation It likes cool and drained soils, acidic and humusy, in the shade. |
Over It occurs where vegetation is dense, such as moist spruce-fir forests, avalanche chutes, along streams, and shrub thickets. |
Shield Ferns, Buckler Ferns, Holly Ferns and their Relatives Propagation: Hardy species by division of crowns in April, also by spores sown on sterilised loam and kept close under glass cover. |
Ferns suitable for Cold-Hardy. Hardy Polystichum fern. Heights vary from 12-36 inches (30-90cms). |
It is found in deep woods in the mountains. Fronds grow to 100 cms (40 inches). It has a conspicuously chaffy fiddlehead and leaf stalk. Pinnae are deeply cut making it appear doubly pinnate. Bulblets form at the base of pinnae near the tip and may grow into a new plant when the frond touches the ground! |
English: Polystichum andersonii in Arboretum Rogow, Poland English: Polystichum andersonii in Arboretum Rogow, Poland |
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Polystichum munitum Hardy in Zone 7 Polystichum means many rows, referring to the arrangement of the spore cases on the undersides of the fronds. Munitum means armed with teeth, referring to its toothed fronds. Western Sword Fern is also known as Sword Holly Fern, Giant Holly Fern, Christmas Fern, Pineland Sword Fern, or Chamisso’s Shield Fern. |
Ascending to erect rhizomes and evergreen fronds.This species does best in moist, cool climates and does not grow well in the eastern of Southeastern United States. The species is native to the western United States, Canada, Alaska (Yukon), and Mexico (Guadalupe Island); it is naturalized in Europe. |
35-47 x 23-47 The dark green fronds of this fern grow in a tight clump spreading out radially from a round base. Individual fronds live for 1.5 to 2.5 years and remain attached to the rhizome after withering. Trim off dead fronds in early spring before new growth begins. |
Shield Ferns, Buckler Ferns, Holly Ferns and their Relatives Propagation: Hardy species by division of crowns in April, also by spores sown on sterilised loam and kept close under glass cover. |
Ferns suitable for Cold-Hardy. Hardy Polystichum ferns. Heights vary from 12-36 inches (30-90cms). |
The fronds are used by florists to include in vases. Hardy fern for a shady, moist situation in the garden or fernery. Plants are quite cold tolerant. The preferred habitat of this fern is the understory of moist coniferous woodlands at low elevations. It grows best in well-drained acidic soil of rich humus and small stones. It is very resilient and survives occasional droughts, but flourishes only with consistent moisture and light sunlight, and it prefers cool weather. |
Phenology: Fronds partially unroll their “fiddleheads” by late May; by late July the spores are near maturity.
Form of |
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Polystichum setiferum (Polystichum angulare, Polypodium angulare, Aspidium angulare) Hardy in Zone 6 This is one of the most graceful of all British native species. |
Erect rhizomes and fronds that are evergreen in warmer climates. Many variants of this species from buds along the rachis - see Section 9 - Propagation . The plants do not like very high humidity. This species is native to Europe. This forms a medium-sized clump of very soft-textured fronds, dark green in colour with a glossy finish. Plants perform best in soils that remain evenly moist, and slightly on the acidic side. |
Height and Spread of |
Shield Ferns, Buckler Ferns, Holly Ferns and their Relatives Propagation: Hardy species by division of crowns in April, also by spores sown on sterilised loam and kept close under glass cover. |
Ferns suitable for Hedge. Hardy Polystichum fern. Heights vary from 12-36 inches (30-90cms). |
It grows in hedge-banks and in lowland woods, preferring, like most of the larger Ferns, the presence of plenty of free (not stagnant) water. Remains evergreen in mild winter regions. Attractive as a specimen, massed, or in containers. |
Buds along the rachis of American Plant Food Company, 7405 River Road, Bethesda MD. Polystichum setiferum . Polystichum setiferum in botanical garden in Batumi |
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Polystichum scopulinum Very Hardy in Zone 4 It is native to much of western North America, and it is known from disjunct occurrences in eastern Canada, as well. |
Ascending to erect rhizomes and leathery, semi-evergreen fronds. This species is difficult to grow. This fern produces several erect, narrowly lance-shaped leaves up to 50 centimeters (20 inches) in length. The leaves narrow near the bases. Each leaf is divided into many lance-shaped or oblong leaflets up to 3 centimeters (1.2 inches) long. The toothed leaflets are sometimes twisted on their axes and overlapping. The species is native to western North America and Canada. |
20 x 12-40
Habitat in Moist rock crevices in subalpine zone, and moist rocks along rivers in the valleys. |
Shield Ferns, Buckler Ferns, Holly Ferns and their Relatives Propagation: Hardy species by division of crowns in April, also by spores sown on sterilised loam and kept close under glass cover. |
Ferns suitable for Cold-Hardy. Hardy Polystichum fern. Heights vary from 12-36 inches (30-90cms). Rock crevices and at base of boulders, serpentine to acidic substrates, usually exposed to full sun; 0--3500 m; B.C., Nfld., Que.; Ariz., Calif., Colo., Idaho, Mont., Nev., Oreg., Utah, Wash., Wyo. |
A small leathery fern suitable for planting among rocks. It is found in dry coniferous forest or more commonly on cliffs and talus slopes. It is more frequent east of the Cascades and the Rocky Mountains; it also grows in eastern Canada. It grows in rocky habitat, often in full sun. It is widespread but mostly found in small populations, and is noted to be most abundant on serpentine soils - Soldiers Delight Natural Environmental Area in Baltimore County, Maryland, covers 1,900 acres of serpentine barren. The area has over 38 rare, threatened, or endangered plant species as well as rare insects, rocks and minerals. |
Fig. 35. Polystichum scopulinum from the second edition of An Illustrated Flora of the Northern United States, Canada and the British Possessions (New York, 1913). By Nathaniel Lord Britton & Addison Brown via Wikimedia Commons Polystichum scopulinum by Sheri Hagwood. Bureau of Land Management. United States, ID, Bureau of Land Management Jarbidge Resource Area. August 1, 2006 via Wikimedia Commons |
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Polystichum triangulum (Polystichum echinatum) Zone 9(10) |
It forms a spreading rosette of narrow, bright green fronds; the pinnae being deltoid to triangular in shape. Native to South America and West Indies. |
Polystichum triangulum (triangular), synonym Polystichum echina-tum, is a small fern that is recommended for Zone 9 and possibly Zone 10. It is most attractive with shiny diamond to triangular-shaped pinnae. If it is not suitable for your garden, try it as a house-plant where, with fronds up to 18 in. (45 cm), it is showy, manageable, and unusual. It can be reproduced by small bulbils on the frond tips. |
Shield Ferns, Buckler Ferns, Holly Ferns and their Relatives Propagation: Greenhouse species by spores sown in sandy peat any time; division in March. |
Ferns suitable for Acid Soil. Greenhouse Polystichum fern. Heights vary from 12-36 inches (30-90cms). |
Although reported as being associated with calcareous rocks, this species grows well in acid, loamy soils. |
If you grow and sell ferns in any country, please tell me so that I can put them on this website and inform others where they can be bought online via mail-order. If you would provide photos and fern details to be only used by me on this website, they would be gratefully received, since I could assume that the photo was a valid one in regard to its name of fern in its filename to that fern in the photo. |
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Ceratopteris thalictroides Tender in Zone 9 It is found throughout the tropics of the world, except Africa. Tropical America - Florida. The name comes from the Greek keras, horn, and pteris, fern, alluding to the antler-like fertile leaves.
AC Tropical Fish (Aquatic Community.com, Aquatic Community) was founded in 2004 as a resource and meeting place for tropical aquarium fish keepers. It has since then grown into a comprehensive aquarium portal featuring information not only about tropical freshwater fish but also about all other types of aquarium fish. |
Buds tend to grow on dying fronds. The bright green, finely-cut, fertile leaves are erect and are produced above water. In Asia the foliage of this species is used in salads and is said to have a peppery taste. This genus has the distinction of having the fastest life cycle of any fern. The plants can take as little as 1 month to go from spores to mature, spore-bearing plants, though 3 or 4 months is more typical. The plants live about 1 year and are usuually perpetuated by the numerous buds that form on the blade surface, typically in margins of the sinuses. The buds detach and float away from the parent plant. It can provide useful shade to shyer fish and small fry. The dense roots are said to take nutrients out of the water helping to prevent the growth of algae. |
30 x |
Aquatic Ferns
Propagation: Mature plants can be propagated by division of the rhizome, which may be allowed to float or, if grown submerged, held in place by coarse gravel. If plants are to be rooted in mud, garden soil with little organic matter will do. Some growers recommend a mixture of about half peat and half sand mixed with 10% top soil. Plants that are rooted in mud need their fronds kept moist. Temperatures must be maintained close to 27C (80F) for good growth. The plants and buds decline when temperatures are below 20C (68F), and if lost, new plants must be started from spores. Propagation: By spores sown in February on surface of compost in pan or water as for culture in next column, vivaparous forms increased by pegging down leaves into soft mud, detaching later. |
Suitable for Aquatic Ferns. Stove aquatic fern that is usually only of annual duration, but when carefully grown may be a biennial. It grows best in soil with a pH reading of 5-9 and in very high amounts of light. It usually grows quickly. |
Grows in wet garden soil. Typically, the plants root in mud. It succeeds best in a pot of good loam, wholly submerged in a tank of warm water in a stove house. It produces spores freely, or may be propagated by the young plants that form on all the fronds. Swampy areas, swamp forests, sago (Metroxylon) swamps, marshes, natural and man-made ponds, mostly in stagnant water bodies or in still pockets along slow flowing rivers, full sun to moderate shade, from sea level to 1300 m, but mostly less than 500 m altitude. Sometimes massed on or around logs or other floating vegetation, once recorded in a fresh-water mangrove (Sonneratia) growing among the finger-like pneumatophores. "The water temperature should ideally be kept above 20 degrees C." from the Aquatic Community. |
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English: Foliage of Ceratopteris thalictroides Figure: Die Pflanzenwelt Afrikas, insbesondere seiner tropischen Gebiete : Grundzge der Pflanzen-verbreitung im Afrika und die Charak-terpflanzen Afrikas. Ceratopteris thalictroides. By Engler, Adolf, 1844-1930 via Wikimedia Commons
English: Two floating aquatic fern: Salvinia minima (right) and Ceratopteris thalictroides (left) floating in aquarium. Identified by user Le.Loup.Gris. |
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Pilularia globulifera Hardy in Zone 5 Native to Europe - found in western Central Europe and scattered throughout the British Isles in shallow water at edges of ponds, ricefields, marshy ground, wet heaths, often submerged, in acid substrata; very local and absent from many counties; local in Ireland. |
Leaves arising from a creeping rhizome with nodes 1-4 cm (0.5-1.5inches) apart, 3-10 (15) (1.2-4 inches) long, subulate. This tiny plant is a type of creeping fern. It is hard to spot because it has thin, grass-like leaves and often grows with water grasses or small rushes. The ‘pills’ are tiny round spore cases at the bases of the stems. It can still be found at a number of sites scattered across Britain, but is internationally threatened, as it is declining across its whole European range. |
About 3 inches (8 cms) tall, which is easily recognised by the characteristically unfurling leaves and the large (3 mm), round sporocarps, if present. Lime green round stem-like leaves or fronds approximately 1-1.5mm diameter. Fronds unfurl from tight coils, and you can often see 1 or 2 fronds which have yet to unfurl even late into the season. Fronds can grow up to 8cm tall, often standing upright from the ground or above the surface of the water, but they can be submerged. The fronds are rarely straight and have a kinky or wavy appearance, especially when young. The fronds arise singly, or at most 3 shoots, from a rhizome (horizontal underground stem), not in clumps or tussocks (as seen in grasses and rushes). |
Aquatic Ferns
Propagation: Pillwort is a specialist of bare pond edge habitats. It is not a good competitor and only thrives where there are few other plants. Like many specialists, it has some key habitat requirements:- |
Suitable for Aquatic Ferns. Stove and Greenhouse evergreen ferns. Pillwort can be grown in a "bog garden" or as a marginal aquatic in a garden pond. |
Grows well under high light in sandy or silty garden soil kept wet. The plants can also grow submerged or partly submerged. Do not let them dry out completely. This speces is a rapid and robust grower in moist to wet soil. It grows at edges of lakes, ponds, ditches and marshes, on wet clay or clay-sand soil (that are submerged for at least part of the year), sometimes in water up to 30 cm (12 in) deep. Some of the plants growing in association with this species in the UK include water celery (Apium inundatum), marsh pennywort (Hydrocotyle vulgaris) and lesser spearwort (Ranunculus flammula). |
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English: Single creeping plant of the fern species Pillwort, Pilularia globulifera, on wet ground. English: Mature sporocarps at the base of the leaves from the fern species Pillwort, Pilularia globulifera. English: Field of the fern species Pillwort, Pilularia globulifera, in a natural habitat (alternating wet pond bottom). Pilularia globulifera - Image:Illustration Pilularia globulifera0.jpg from Prof. Dr. Otto Wilhelm Thomé Flora von Deutschland, Österreich und der Schweiz 1885, Gera, Germany via Wikimedia Commons |
Pilularia are small, sedge-like or grass-like plants. They can be distinguished from grasses and sedges by the coiled tips of their young leaves. Of little ornamental value, this genus is best used as part of a small, aquatic dish-garden or in bog or marsh plantings. |
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Nephrolepis acuminata (Nephrolepis davallioides, Nephrolepis biserrata and other synonyms) Malaysia, Indonesia Tropical - Semi-tropical The genus name comes from the Greek nephros, kidney, and lepis, scale, referrring to the kidney-bean-shaped indusia. |
The fertile pinnae are deeply lobed and attractive. Tufted clusters of golden-yellow foliage arising from underground rhizomes. The round sori (clusters of spore-bearing organs) are in two rows near the margins on the underside of the pinnae. The fronds grow upright at first, then arch gracefully downwards. They grow in lovely arching rosette shaped and spread by runners. |
24-48 x 2-6 |
Boston Ferns (Nephrolepis)
Propagation: By spores sown on surface of pans of sandy peat under bell-glass and placed in temperature 75-85F (24-30C) any time; division of plants, February-April; or by pegging down creeping stems bearing young plants and removing when rooted. The most common problem in caring for established plants is overwatering combined with poor drainage. These ferns generally tolerate short periods of dryness. |
Suitable for Conservatory or Heated Greenhouse. Basket Fern. Stove Evergreen Ferns. Fronds linear, narrow, once divided, plain or crested. First introduced late eighteenth century. |
This fern can be grown as a garden plant in the tropics, as well as in a large container. |
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Nephrolepis biserrata (Nephrolepis acuta, Aspidium biserrata) Malaysia, Malesiana, Indonesia, Southeast Asia, Australia, North and South America, the Pacific islands, Native Plants of South Florida Tropical - Semi-tropical
The genus name comes from the Greek nephros, kidney, and lepis, scale, referrring to the kidney-bean-shaped indusia. Species biserrata means double-toothed and refers to the leaf margin of the fern. |
The fertile pinnae are deeply lobed and attractive. Tufted clusters of golden-yellow foliage arising from underground rhizomes. The round sori (clusters of spore-bearing organs) are in two rows near the margins on the underside of the pinnae. The fronds grow upright at first, then arch gracefully downwards. They grow in lovely arching rosette shaped and spread by runners. Grow in Part Shade. |
24-48 x 2-6 |
Boston Ferns (Nephrolepis)
Propagation: By spores sown on surface of pans of sandy peat under bell-glass and placed in temperature 75-85F (24-30C) any time; division of plants, February-April; or by pegging down creeping stems bearing young plants and removing when rooted. The most common problem in caring for established plants is overwatering combined with poor drainage. These ferns generally tolerate short periods of dryness. |
Suitable for Conservatory or Heated Greenhouse. Basket Fern. Stove Evergreen Ferns. Fronds linear, narrow, once divided, plain or crested. First introduced late eighteenth century. |
This fern can be grown as a garden plant in the tropics, as well as in a large container. Found in swamps and hammocks. |
Nephrolepis biserrata Nephrolepis biserrata
Further photos. |
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Nephrolepis cordifolia (Nephrolepis tuberosa, Polypodium cordifolium, Aspidium cordifolium) Native to northern Australia and Asia. The genus name comes from the Greek nephros, kidney, and lepis, scale, referrring to the kidney-bean-shaped indusia. Nephrolepis cordifolia has become an invasive species is some areas where it has been introduced. In New Zealand it is listed on the National Pest Plant Accord, which prohibits the sale, cultivation and distribution of the plant. |
Nephrolepis cordifolia is a wood fern that typically grows in woodland areas. Both fertile and sterile fronds are pinnate, up to 3 feet in length and 3 inches wide. There are many leaflets, or pinnae, ranging from 40-100 mm (1.5 to 4 inches) on each side of the rachis. Each pinna is oblong to lanceolate with an auricle that overlaps rachis. Rhizomes are orange/brown to pale brown with linear scales having hair like tips. Stolons are straw colored and produce small underground tubers. The presence of tubers distinguishes sword fern from the native Nephrolepis exaltata fern. Numerous sori (spore containing structures) are also produced between the leaflet midvein and margin. Dispersal occurs via spores and through the movement of stolons, tubers, and rhizomes.
Hardy to 25°F. |
24-36 x 24-36 |
Boston Ferns (Nephrolepis)
Propagation: By spores sown on surface of pans of sandy peat under bell-glass and placed in temperature 75-85F (24-30C) any time; division of plants, February-April; or by pegging down creeping stems bearing young plants and removing when rooted. The most common problem in caring for established plants is overwatering combined with poor drainage. These ferns generally tolerate short periods of dryness. Grows in wet, shady places, limestone ledges, cliffs, rock and roadsides in North America. |
Suitable for Basket fern. Stove Evergreen Ferns. Fronds linear, narrow, once divided, plain or crested. First introduced late eighteenth century. |
Ground cover in tropical and subtropical areas. Grows from shade to full sun (grows in full sun if given ample water) in soil, among rocks or as an epiphyte (particularly on palm trunks). It is colony former and is popularly grown in temperate regions but in the tropics is generally regarded as a weed. It can be grown in gardens, pots or baskets. Bayabang grows in the Philippines as a hedge plant.
Also in |
Español: Cola de Quetzal (Nephrolepis cordifolia), jardín botánico de Tallinn, Estonia
Nephrolepis cordifolia (L.) C.Presl from Rodney Ecological District. This image has been released as "CCBY" by Auckland War Memorial Museum. By Ewen Cameron via Wikimedia Commons.
Nephrolepis cordifolia - Sori. Date 19 March 2008. By Ixitixel via Wikimedia Commons |
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Nephrolepis exaltata Florida, Mexico, Brazil, West Indies. Tender to Semi-tender in USDA Zones 9-11. |
The fronds of Nephrolepis exaltata are 50–250 centimetres (20–98 in) long and 6–15 centimetres (2.4–5.9 in) broad, with alternate pinnae (the small "leaflets" on either side of the midrib), each pinna being 2–8 centimetres (0.79–3.15 in) long. Although the fern may appear totally dead due to frost, it will re-emerge in the spring. In general, the Boston fern likes damp, but not soggy soil that is rich in nutrients. Of the common cultivated ferns, the Boston fern is the most tolerant to drought. The fern thrives best in humid conditions, so when grown as a house plant it becomes necessary to mist the plant when relative humidity falls below around 80%. Although outdoors this plant prefers partial shade or full shade, inside it doesn't grow in shade and feels best in bright filtered light. |
20-100 x 48 It is safe for pets as it is known to be non-toxic. In 1989, the NASA Clean Air Study showed that the Boston Fern could filter Formaldehyde, Xylene and Toluene from the air.
Nephrolepis exaltata is classified as an invasive alien plant in South Africa. In some provinces it must, by law, be eradicated. |
Boston Ferns (Nephrolepis)
Propagation: By spores sown on surface of pans of sandy peat under bell-glass and placed in temperature 75-85F (24-30C) any time; division of plants, February-April; or by pegging down creeping stems bearing young plants and removing when rooted. This plant is usually propagated by division of the rooted runners, as named cultivars will not produce true spores. The most common problem in caring for established plants is overwatering combined with poor drainage. These ferns generally tolerate short periods of dryness. |
Suitable for Conservatory or Warm Greenhouse. Stove Evergreen Ferns. Fronds linear, narrow, once divided, plain or crested. First introduced late eighteenth century. It is a very popular house plant, often grown in hanging baskets. Grow in steamy bathroom. |
A medium-large fern with short, erect, stolonoiferous rhizomes. Grows best under medium to high light in moist-dry potting mixes. So commonly grown are these ferns that commercial growers have established the precise cultural conditions for optimum growth: 2000 foot-candles of light, temperatures between 18 and 35 degrees Centigrade (65-95 degrees Fahrenheit), fertilizers with a 3-1-2 composition, micronutrients, and dolomite to adjust the pH to 5.5. Use a mix with good drainage, and water when the soil surface is slightly dry. To prevent breakage and decay of fronds, water should be applied at the soil level, not on the foliage. Poor foliage results when temperatures are below 18 degrees Centigrade (65 Fahrenheit) for more than a month. Repotting just before the active growth in spring is recommended. |
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Nephrolepis exaltata (in a greenhouse). English: Greenhouses of the Botanical garden (Saint Petersburg).
The Boston Sword Fern - Nephrolepis exaltata is widely planted from an origin in Central America and the Caribbean. Lomariopsidaceae Lotusland, Montecito, California. By Dick Culbert from Gibsons, B.C., Canada via Wikimedia Commons.
Nephrolepis exaltata. I take this photo from Chinese Medicine Garden in Saikung Hong Kong on 19th March 2006. By Xiaowei (talk | contribs) via Wikimedia Commons. |
See more photos. |
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Nephrolepis pendula (Nephrolepis cordifolia) Native to tropical America, Mexico, Central and South America Tender to Semi-tender in Zones 9a-11 The genus name comes from the Greek nephros, kidney, and lepis, scale, referrring to the kidney-bean-shaped indusia. |
This fern is ginormous! It commonly grows up to 6′ tall, and its fronds can grow to 20 feet long. That may not work great for small apartments in the city, but they look fantastic in hotels, interior atriums and malls where they can stretch to their full glory. |
36-72 x 24-36 |
Boston Ferns (Nephrolepis)
Propagation: By spores sown on surface of pans of sandy peat under bell-glass and placed in temperature 75-85F (24-30C) any time; division of plants, February-April; or by pegging down creeping stems bearing young plants and removing when rooted. The most common problem in caring for established plants is overwatering combined with poor drainage. These ferns generally tolerate short periods of dryness. |
Suitable for Basket Fern with Pendulous Fronds for Conservatory. Stove Evergreen Ferns. Fronds linear, narrow, once divided, plain or crested. First introduced late eighteenth century. |
A large evergreen fern with short, stoloniferous rhizomes.Grows well under medium light in moist potting mix. It is a choice fern for hanging baskets becase of its long, narrow, pendent fronds. Prefers part shade and relatively high humidity to grow well. |
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Cheilanthes austrotenuifolia (Cheilanthes tenuifolia) Australia, New Zealand The Seeds of South Australia database contains 143 families, 837 genera and 3,103 native species with 29,293 images (also with 400 introduced species). |
Grow in a terrarium set up with a stony potting mix and kept on the dry side (by sparse watering and leaving the lid off for long periods), Cheilanthes can sometimes be grown to perfection. They may need brighter light than other ferns usually grown in terrariums. Bright green fronds grow from an underground rhizome. Fronds die down in summer and return with the rain in Autumn. Spore cases partially enclosed by scalloped margins of lobes. |
4-20 x Fronds crowded, mostly 15–30 cm high, 3–10 cm wide at the widest point; stipe red-brown, dark brown or black, covered densely at the base with transparent scales; lamina lanceolate to triangular, 2–3-pinnate, glabrous above, sparsely scaly below; ultimate segments sessile, 2–6 mm long and 1–3 mm wide, with crenate margins. |
Cloak, Lip, Hand Ferns and Their Hardy Relatives (Bommeria, Cheilanthes, Doryopteris, Gymnopteris, Hemionitis, Notholaena, Paraceterach, Pellae, Pleurosorus, Quercifilix)
Propagation: By spores sown on fine sandy peat, kept moist and shaded under bell-glass. |
Suitable for Conservatory and Heated Greenhouse. Stove and Greenhouse Ferns. First introduced late eighteenth century. |
This is a dwarf fern which develops into compact clumps of bright green, finely divided fronds. It usually occurs in rocky situations and plants look particularly appealing in a rock garden, especially when situated against dark rocks. Plants require well-drained, acid, humus-rich soils in a sunny situation. Once a common understory plant along streams in moist areas of Southern Australia. Grows in rocky ground in open forest or on exposed rocky slopes in New South Wales. Garden Use in Victoria, Australia:While difficult to establish it is very tough for rockeries and exposed positions provided it has root protection. |
Cheilanthes austrotenuifolia H.M.Quirk & T.C.Chambers, Black Mountain, Canberra, ACT, 4 November 2010. By Donald Hobern from Copenhagen, Denmark via Wikimedia Commons. See more photos from Seeds of South Australia. |
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Cheilanthes californica (Hypolepis californica, Adiantopsis californica) California, Mexico Needs dry atmosphere. |
Grow in a terrarium set up with a stony potting mix and kept on the dry side (by sparse watering and leaving the lid off for long periods), Cheilanthes can sometimes be grown to perfection. They may need brighter light than other ferns usually grown in terrariums. It has lacy fronds of an attractive fresh green. Aspidotis californica has leaves that are thin and dissected into many triangular leaflets which are subdivided into small segments with curled teeth. |
2-6 x 2-4 |
Cloak, Lip, Hand Ferns and Their Hardy Relatives (Bommeria, Cheilanthes, Doryopteris, Gymnopteris, Hemionitis, Notholaena, Paraceterach, Pellae, Pleurosorus, Quercifilix)
Propagation: By spores sown on fine sandy peat, kept moist and shaded under bell-glass. |
Suitable for Rock Garden and Wall Fern. Stove and Greenhouse Ferns. First introduced late eighteenth century. |
Greenhouse Fern. It occurs naturally on shaded, rock sites. In cultivation, it has proved tricky to maintain requiring a very porous, acid mixture, bright light but not sun, and ample air movement. Plants are susceptible to overwatering. It grows in rock cracks and crevices in many types of habitat, including Chaparral, Yellow pine forest, Foothill oak woodland, and Valley grassland in California, USA. |
See Botanical Figure. |
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Cheilanthes distans + Australia, New Zealand, Lord Howe Island, New Caledonia This small plant is native to various parts of Australia including Western Australia and New South Wales in the east. It is occasionally seen around Sydney in rocky exposed areas. The Bristly Cloak Fern grows in areas of high rainfall as well as the semi arid areas of inland Australia. distans: distant (widely spaced female flowers). Found in drier, exposed areas in the eastern parts of the North Island from the Bay of Islands the middle of the South Island of New Zealand. |
Grow in a terrarium set up with a stony potting mix and kept on the dry side (by sparse watering and leaving the lid off for long periods), Cheilanthes can sometimes be grown to perfection. They may need brighter light than other ferns usually grown in terrariums. Fronds crowded, to 30 cm high, < 3 cm wide; stipe red-brown to dark brown, shiny, moderately to densely covered with lanceolate, entire brown scales; lamina narrow-lanceolate, 2-pinnate, upper surface sparsely hairy with simple whitish hairs, lower surface with scales only, golden-brown; ultimate segments sessile, 1–6 mm long and 1 mm wide. |
4-6 x |
Cloak, Lip, Hand Ferns and Their Hardy Relatives (Bommeria, Cheilanthes, Doryopteris, Gymnopteris, Hemionitis, Notholaena, Paraceterach, Pellae, Pleurosorus, Quercifilix)
Propagation: By spores sown on fine sandy peat, kept moist and shaded under bell-glass. Minute spores are wind dispersed (Thorsen et al., 2009). |
Suitable for Rock Garden and Wall Fern. Stove and Greenhouse Ferns. First introduced late eighteenth century. |
A fairly easily grown species which forms clumps of slender fronds that are covered with bristly scales. Attractive in a pot or rock garden. Needs acid, humus-rich soil in a part shade situation. It has a woolly appearance with small white hairs on the top side of the fronds, and a rusty brown underneath. Grows on rocky hillsides in woodland or open forest in New South Wales, Australia. Coastal to montane in dry, rocky habitats with only sparse or no vegetation cover. Often found growing with Asplenium flabellifolium, Cheilanthes sieberi subsp. sieberi and Pellaea calidirupium. More common in the drier eastern parts of the country of New Zealand. |
Cheilanthes distans at UC Berkeley Botanical Garden, California, USA. Identified by sign. Date 31 March 2010. By Stickpen via Wikimedia Commons. See other photos and Noosa's Publications. |
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If you grow and sell ferns in any country, please tell me so that I can put them on this website and inform others where they can be bought online via mail-order. If you would provide photos and fern details to be only used by me on this website, they would be gratefully received, since I could assume that the photo was a valid one in regard to its name of fern in its filename to that fern in the photo. |
Site design and content copyright ©January 2009. |
Fern Grower's Manual by Barbara Joe Hoshizaki & Robbin C. Moran. Revised and Expanded Edition. Published in 2001 by Timber Press, Inc. Reprinted 2002, 2006. ISBN-13:978-0-88192-495-4. |
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USE OF FERN WITH PHOTOS
using information from Fern Grower's Manual by Barbara Joe Hoshizaki & Robbin C. Moran and
The Encyclopaedia of Ferns An Introduction to Ferns, their Structure, Biology, Economic Importance, Cultivation and Propagation by David L. Jones ISBN 0 88192 054 1
Outdoor Use in
Northeastern United States Zones 3-6
Southeastern United States Zones 6-8
Southern Florida and Hawaii Zones 10-11
Central United States Zones 3-6
Northwestern United States Zones 5-8 with some Zone 9
Southwestern United States Zones 6-9
Coastal Central and Southern California Zones 9-10
Accent
Aquatic 1, 2
Basket 1,
Ferns for Hanging Baskets 2, 3, 4, 5, 6
Ferns for Hanging Baskets with Pendulous Fronds or weeping Growth Habit 7, 8
Bog or Wet-Soil 1,
Ferns for Wet Soils 2, 3
Border and Foundation 1, 2
Grow in Coastal Region
Cold-hardy Ferns 1, 2, 3, 4, 5, 6
Colour in Fern Fronds 1, 2, 3, 4
Conservatory (Stove House) or Heated Greenhouse 1, 2, 3, 4, 5
Drier Soil 1, 2, 3, 4
Grows on Rock (epilithic) 1, 2
Borne on Leaf (epiphyllous) 1, 2
Grows on another Plant (epiphyte) 1, 2
Evergreen and Deciduous
Fronds in Floral Decorations
Ferns for Acid Soil 1,
Lime-hating (Calcifluges) 2, 3, 4, 5
Ferns for Basic or Limestone Soil 1,
Ferns Found on Limestone or Basic Soils (Calciphiles) 2, 3, 4, 5, 6, 7, 8, 9, 10
Ferns for Ground Cover 1,
Ground Cover Ferns 2, 3, 4, 5
Ferns of the Atlantic Fringe with associated plants (1 - Atlantic Cliff-top Grassland, Ledges and Rough Slopes; 2 - Clay Coasts and Dunes of South-East Ireland; 3 - Limestones of Western Atlantic Coasts; 4 - Hebridean Machair; 5 - Horsetail Flushes, Ditches and Stream Margins; 6 - Water Margin Osmunda Habitats; 7 - Western, Low-lying, Wet, Acid Woodlands; 8 - Western, Oak and Oak-Birch Woodlands and Ravines, in the UK and Ireland)
Ferns in Coastal District with associated plants (Hard Rock Cliffs, Soft Rock Cliffs, Clay Coasts, or Coastal Sand-Dunes in the UK)
Ferns of Grasslands and Rock Outcrops (Grasslands; Rocks, Quarries and Mines in the UK)
Ferns of Heath and Moorland with associated plants (1 - Bracken Heath; 2 - Ferns of Moist Heathland Slopes and Margins of Rills and Streams; 3 - Heathland Horsetails, 4 - Heathland Clubmosses, in the UK)
Ferns of Lower Mountain Habitats with associated plants (1 - Upland Slopes and Screes; 2 - Base-rich, Upland Springs and Flushes; 3 - Base-rich, Upland, Streamside Sands and Gravels; 4 - Juniper Shrub Woodland, in the UK)
Ferns for Man-Made Landscapes with associated plants (South-western Hedgebanks, Hedgerows and Ditches, Walls and Stonework, Water Mills and Wells, Lime Kilns and abandoned Lime-Workings, Pit heaps and Shale Bings, Canals, Railways and Their Environs in the UK)
Ferns of Upper Mountain Habitats with associated plants (1 - High Mountain, Basic Cliffs and Ledges; 2 - High, Cliff Gullies; 3 - High Mountain Corries, Snow Patches and Fern beds; 4 - Ridges, Plateaux and High Summits, in the UK)
Ferns for Wetlands with associated plants (1- Ponds, Flooded Mineral Workings and Wet Heathland Hollows; 2 - Lakes and Reservoirs; 3 - Fens; 4 - Ferns of the Norfolk Broads' Fens; 5 - Willow Epiphytes in the UK)
Ferns in Woodland with associated plants (1 - Dry, Lowland, Deciduous Woodland; 2 - Inland, Limestone, Valley Woodland; 3 - Base-rich Clay, Valley Woodland; 4 - Basic, Spring-fed Woodland; 5 - Ravine Woodland on Mixed Rock-types; 6 - Native Pine Forest in the UK)
Ferns in Hedges or Hedgebanks
Outdoor Containers 1, 2, 3, 4, 5, 6, 7
Rapidly Growing Fern 1, 2
Resurrection Fern
Rock Garden and Wall Ferns 1, 2, 3, 4, 5
Shade Tolerant 1, 2, 3, 4
Slowly Growing Fern
Sun Tolerant 1, 2, 3, 4
House Fern in Trough Garden 1,
Fern Suitable for
Indoor Decoration 2, 3, 4, 5, 6
House Fern in Terrarium, Wardian Case or
Bottle Garden 1,
Ferns suitable for Terrariums, Wardian Cases 2, 3, 4,
5, 6
Grow in Woodlands 1, 2, 3, 4
TYPE OF FERN TO GROW WITH PHOTOS
using information from
Fern Grower's Manual by Barbara Joe Hoshizaki & Robbin C. Moran and
The Encyclopaedia of Ferns An Introduction to Ferns, their Structure, Biology, Economic Importance, Cultivation and Propagation by David L. Jones ISBN 0 88192 054 1
Aquatic Ferns (Azolla, Ceratopteris, Marsilea, Pilularia, Regnellidium, Salvinia)
Boston ferns (Nephrolepis exaltata), Fishbone ferns (Nephrolepis cordifolia), Lace ferns and Sword ferns
Cloak, Lip, Hand Ferns and their Hardy Relatives (Bommeria, Cheilanthes, Doryopteris, Gymnopteris, Hemionitis, Notholaena, Paraceterach, Pellae, Pleurosorus, Quercifilix) 1,
2, 3
Davallia Ferns (Araiostegia, Davallia, Davallodes, Gymno-grammitis, Humata, Leucostegia, Scyphularia, Trogostolon) 1, 2
Fern Allies (Psilotums or Whisk Ferns, Lycopodiums or Ground Pines, Selaginellas or Spike Mosses, and Equisetums, Horsetails or Scouring Rushes) 1, 2
Filmy and Crepe Ferns (Hymenophyllum, Trichomanes, Leptopteris) 1, 2
Lacy Ground Ferns (Culcita, Dennstaedtia, Histiopteris, Hypolepis, Leptolepia, Microlepia, Paesia, Pteridium) 1, 2
Lady Ferns and Their Allies (Allantodia, Athyrium, Diplazium, Lunathyrium, Pseudo-cystopteris, Callipteris, Cornopteris, Cystopteris) 1, 2
Maidenhair Ferns (Adiantum) 1, 2
Miscellaneous Ferns (Acrostichum, Actiniopteris, Anemia, Anogramma, Anopteris, Blotiella, Bolbitis, Christella, Coniogramma, Cryptogramma, Ctenitis, Cyclosorus, Didymochlaena, Dipteris, Elaphoglossum, Equisetum, Gymnocarpium, Llavea, Lonchitis, Lygodium, Macrothelypteris, Oeontrichia, Oleandra, Onoclea, Onychium, Oreopteris, Parathelypteris, Phegopteris, Photinopteris, Pityrogramma, Pneumatopteris, Psilotum, Stenochlaena, Thelypteris, Vittaria)
1, 2, 3, 4 including Fern Allies of Equisetum and Psilotum or Whisk Ferns
Polypodium Ferns and Relatives (Anarthropteris, Belvisia, Campyloneurum, Colysis, Crypsinus, Dictymia, Gonphlebium, Lecanopteris, Lemmaphyllum, Lexogramme, Microgramma, Microsorum, Niphidium, Phlebodium, Phymatosurus, Pleopeltis, Polypodium, Pyrrosia, Selliguea) 1, 2, 3
Primitive Ferns and Fern Oddities (Angiopteris, Botrychium, Christensenia, Danaea, Helminthostachys, Marattia, Ophioglossum, Osmunda and Todea)
Scrambling, Umbrella, Coral and Pouch Ferns (Dicranopteris, Diploptergium, Gleichenia, Sticherus)
Shield, Buckler, Holly Ferns and their Relatives (Arachniodes, Cyrtomium, Dryopteris, Lastreopsis, Matteuccia, Polystichum, Rumohra, Tectaria and Woodsia) 1, 2, 3, 4
Spleenworts Ferns (Asplenium) 1, 2, 3
Staghorns, Elkhorns and other large epiphytes (Aglaomorpha, Drynaria, Merinthosorus, Platycerium, Pseudodrynaria) 1, 2
Fern Allies - Tassel Ferns and Clubmosses (Lycopodium)
The Brakes (Pteris) 1, 2
Tree Ferns (Cibotium, Cnemidaria, Cyathea, Dicksonia, Nephelea and Trichipteris) 1, 2
Water, Hard, Rasp and Chain Ferns (Blechnum, Doodia, Woodwardia, Sadleria) 1, 2
Xerophytic Ferns (Actinopteris, Astrolepis, Cheilanthes, Doryopteris, Notholaena, Pellaea, Pityrogramma) 1, 2